We have the 2009 bee, (little one born a tad too early for the bee plus) and the cameleon. I love the cameleon for the first 6 months as the carry cot is so much like a moses basket, the seat unit is also very sturdy as they get bigger, however found the pram a bit of hardwork on public transport as the back wheels stick out a little bit, so for example on the bus its a bit of a squeeze if there is already a pram in the buggy bay. The cameleon also folds down to quite a large size. OH hates how big it is and refuses to go shopping with it as it leaves little space for the shopping, he hates that it folds in two parts. The bee is much more compact and ideal for hopping on and off the bus and tube, it is lightweight and can easily be carried up and down stairs even with a toddler in it, whereas the cam is heavier with toddler in. It folds smaller and in one part which is much easier for when it does need to be put down. We had the cocoon and this lasted us the whole of our first winter, little one also slept soundly in this and felt she was much more secure in this as she was strapped in. when she outgrew this i brought a liner as i felt the bee seat looked a little too uncomfortable without any extra support. The one thing which annoyed me with this bee is that by no fault of its own, when the bee plus came out, bugaboo made all these new coloured hoods that didn't fit our model, so we was stuck with the 6 original colours to choose from. The footmuff for us was a great investment as it fitted both models. The carseat adaptors were a must for moving sleeping baby from car to pram without waking her. All in all I would say the cameleon is fab if you do lots for walking and it glides along different surfaces a lot smoother than the bee, and is also better at being bumped down stairs as the suspension is alot better, The bee is better if you are on and off public transport or in and out the car, the seat unit is a little more fiddley the turn from parent to outward facing than the cameleon but you get the hang of it the more you do it. Either model will not let you down, good luck with your choice x